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Hearing (Sense) Information

Hearing (or audition; adjectival form: "auditory" or "aural") is the ability to perceive sound by detecting vibrations through an organ such as the ear.[1] It is one of the traditional five senses. The inability to hear is called deafness.

In humans and other vertebrates, hearing is performed primarily by the auditory system: vibrations are detected by the ear and transduced into nerve impulses that are perceived by the brain (primarily in the temporal lobe). Like touch, audition requires sensitivity to the movement of molecules in the world outside the organism. Both hearing and touch are types of mechanosensation.[2]

Contents

Hearing mechanism

The eardrum of an ear simplifies incoming air pressure waves to a single channel of amplitude. In the inner ear, the distribution of vibrations along the length of the basilar membrane is detected by hair cells. The location and intensity of vibrations in the basilar membrane is transmitted to the brain through the auditory nerve.

Hearing tests

Main article: Hearing test

Hearing can be measured by behavioral tests using an audiometer. Electrophysiological tests of hearing can provide accurate measurements of hearing thresholds even in unconscious subjects. Such tests include auditory brainstem evoked potentials (ABR), otoacoustic emissions (OAE) and electrocochleography (EchoG). Technical advances in these tests have allowed hearing screening for infants to become widespread.

Defense mechanism

The hearing structures of many species have defense mechanisms against injury. For example, the muscles of the middle ear (e.g. the tensor tympani muscle) in many mammals contract reflexively in reaction to loud sounds which may injure the hearing ability of the organism.

The idea that the acoustic reflex is a defense mechanism has been a topic for debate in recent years. Commonly cited counterarguments include:

Hearing underwater

Hearing threshold and the ability to localize sound sources are reduced underwater, in which the speed of sound is faster than in air. Underwater hearing is by bone conduction, and localization of sound appears to depend on differences in amplitude detected by bone conduction.[3]

Hearing in animals

Not all sounds are normally audible to all animals. Each species has a range of normal hearing for both loudness (amplitude) and pitch (frequency). Many animals use sound to communicate with each other, and hearing in these species is particularly important for survival and reproduction. In species that use sound as a primary means of communication, hearing is typically most acute for the range of pitches produced in calls and speech.

Frequencies capable of being heard by humans are called audio or sonic. The range is typically considered to be between 20 Hz and 20,000 Hz.[4] Frequencies higher than audio are referred to as ultrasonic, while frequencies below audio are referred to as infrasonic. Some bats use ultrasound for echolocation while in flight. Dogs are able to hear ultrasound, which is the principle of 'silent' dog whistles. Snakes sense infrasound through their bellies, and whales, giraffes, dolphins and elephants use it for communication.

Certain animals have more sensitive hearing than humans which enables them to hear sounds too faint to be detected by humans.

Mathematics

The eardrum of an ear simplifies incoming air pressure waves to a single channel of amplitude. The basilar membrane of the inner ear separates out different frequencies: high frequencies produce a large vibration at the end near the middle ear, and low frequencies a large vibration at the distant end. Thus the ear performs a frequency analysis, roughly similar to a Fourier transform.[5][6] However, the nerve pulses delivered to the brain contain both place and rate information, so the similarity is not strong.

See also

References

  1. ^ Jan Schnupp, Israel Nelken and Andrew King (2011). Auditory Neuroscience. MIT Press. ISBN 026211318X. https://mustelid.physiol.ox.ac.uk/drupal.
  2. ^ Kung C. (2005-08-04). "A possible unifying principle for mechanosensation". Nature 436 (7051): 647–654. doi:10.1038/nature03896. PMID 16079835. http://www.nature.com/nature/journal/v436/n7051/full/nature03896.html.
  3. ^ Shupak A. Sharoni Z. Yanir Y. Keynan Y. Alfie Y. Halpern P. (January 2005). "Underwater Hearing and Sound Localization with and without an Air Interface". Otology & Neurotology 26 (1): 127–130. doi:10.1097/00129492-200501000-00023. http://otology-neurotology.com/pt/re/otoneuroto/abstract.00129492-200501000-00023.htm;jsessionid=Hn3GlTRJcB530CTrCxLlgrJLhv6WyCvpgcBmC0FLJCLWgY5yckpm!1138671057!181195629!8091!-1?index=1&database=ppvovft&results=1&count=10&searchid=1&nav=search.
  4. ^ "Frequency Range of Human Hearing". The Physics Factbook. http://hypertextbook.com/facts/2003/ChrisDAmbrose.shtml.
  5. ^ Deutsch, Diana (1999). The psychology of music. Gulf Professional Publishing. p. 153. ISBN 9780122135651. http://books.google.com/books?id=A3jkobk4yMMC&pg=PA153. Retrieved 24 May 2011.
  6. ^ Hauser, Marc D. (1998). The evolution of communication. MIT Press. p. 190. ISBN 9780262581554. http://books.google.com/books?id=QbunyscCJBoC&pg=PA190. Retrieved 24 May 2011.

Further reading

External links

Wikiquote has a collection of quotations related to: Hearing
Look up hearing in Wiktionary, the free dictionary.
Nervous system: Sensory systems / senses (TA A15)
Special senses

Visual system/sight

Auditory system/hearing

Chemoreception (Olfactory system/smellGustatory system/taste)
Touch Pain (Nociception) • Heat (Thermoception) • Balance (Equilibrioception) • Mechanoreception (Pressure, vibration, proprioception)
Other Sensory receptor
Sensory system: Auditory and Vestibular systems (TA A15.3, TH 3.11.09, GA 10.1029)
Outer ear

Pinna (Helix, Antihelix, Tragus, Antitragus, Incisura anterior auris, Earlobe) • Ear canalAuricular muscles

Eardrum (Umbo, Pars flaccida)
Middle ear
Tympanic cavity

Labyrinthine wall/medial: Oval window · Round windowSecondary tympanic membrane • Prominence of facial canal • Promontory of tympanic cavity

Membranous wall/lateral

Mastoid wall/posterior: Mastoid cellsAditus to mastoid antrumPyramidal eminence

Carotid wall/anterior

Tegmental wall/roof: Epitympanic recess

Jugular wall/floor
Ossicles Malleus (Neck of malleus, Superior ligament of malleus, Lateral ligament of malleus, Anterior ligament of malleus) · Incus (Superior ligament of incus, Posterior ligament of incus) · Stapes (Anular ligament of stapes)
Muscles Stapedius · Tensor tympani
Eustachian tube Bony part of pharyngotympanic tube · Cartilage of pharyngotympanic tube (Torus tubarius)
Inner ear/ (membranous labyrinth, bony labyrinth)
Auditory system Cochlear labyrinth
General cochlea Scala vestibuliHelicotremaScala tympaniModiolusCochlear cupula
Perilymphatic space PerilymphCochlear aqueduct
Cochlear duct / scala media

Reissner's/vestibular membraneBasilar membrane

EndolymphStria vascularisSpiral ligament

Organ of Corti: StereociliaTectorial membraneSulcus spiralis (externus, internus) • Spiral limbus
Cells Claudius cellBoettcher cell
Vestibular system/ Vestibular labyrinth

Static/translations/vestibule/endolymphatic duct: Utricle (Macula) · Saccule (Macula, Endolymphatic sac) · Kinocilium · OtolithVestibular aqueductCanalis reuniens

Kinetic/rotations: Semicircular canals (Superior, Posterior, Horizontal) • Ampullary cupulaAmpullae (Crista ampullaris)

: EAR

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